Richard E. Lenski, Charles Ofria, Robert T. Pennock, Christoph Adami, "The evolutionary origin of complex features" (2003)
"Nature" 8 May 2003, vol. 423, s. 139-144; http://myxo.css.msu.edu/papers/nature2003/Nature03_Complex.pdf --- Patrz też Lenski et al., "The evolutionary origin of complex features. Supplementary information".